1975: PhD in Pharmacology at the University of Hokkaido
1977-1978: Fellow of Humboldt Foundation at the University of Bonn
1979-1980: Fellow of British Counsil at the London Imperial College
1992-1998: Professorship at the University of Nagoya, Gradduate School of Human Informatics
1998-2011: Professorship at the University of Hokkaido, Research Institute for Electronic Science
2005: Nikon Bioimagine Centre
from 2011: Professor emeritus at the University of Hokkaido